YOKOHAMA, Japan – Portland, Maine, native Ian Crocker touched out one of the top butterfly swimmers in the world, winning a gold medal in the 100-meter butterfly during the final day of the Pan Pacific swimming championships Thursday.

Crocker’s gold was one of five first-place finishes Thursday for the United States, which also capped a dominating performance in the meet with a world record in the men’s 4×100-meter medley relay.

Crocker took the 100 butterfly title in 52.45 seconds, ahead of favorite Geoff Huegill of Australia (52.48). Huegill is ranked second in the world in the 100 fly.

Canada’s Mike Mintenko touched in 52.69 for bronze. Mintenko set the early pace at 24.43, with Crocker turning in third place at 24.61, behind Australia’s Adam Pine (24.55).

“I was expecting to be out faster than I was in the prelims and semis, and I was, but I didn’t feel quite as relaxed,” Crocker told USA Swimming. “I’m happy with the result, but I would have liked to have gone faster.”

Crocker, who is a Cheverus High of Portland graduate and will be a junior at the University of Texas this fall, set the 100 fly meet record during a semifinal swim Wednesday. He held the American record until it was broken earlier this month.

Crocker won an Olympic gold medal in 2000 for swimming on a relay.

In the men’s medley relay Thursday, Aaron Peirsol, Brendan Hansen, Michael Phelps and Jason Lezak clocked 3 minutes, 33.48 seconds, eclipsing the previous world mark of 3:33.73 set by the U.S. at the 2000 Olympics in Sydney.

Ian Thorpe anchored the Australian team to second place, adding a silver to his five gold medals at the meet.

“Going into the race we all wanted that record,” said Phelps, who swam the butterfly leg after winning the 200 individual medley earlier in the competition. “We tried for it and were successful.”

The relay victory gave the United States 21 gold medals in the six-day meet. Australia was next with 11.

Natalie Coughlin won the women’s 100 freestyle, collecting her fourth gold medal with a championship record 53.99 seconds. Australia’s Jodie Henry (54.55) finished second and veteran American Jenny Thompson (54.75) was third.

Coughlin, who also won the women’s 100 butterfly, 100 backstroke and was a member of the gold-medal winning 4×200 freestyle team, said she was surprised with her time.

In the 200 IM, Phelps swam a meet record 1:59.70 to edge Japan’s Takahiro Mori (2:00.61) and fellow American Thomas Wilkins (2:01.17). Margaret Hoelzer clocked 2:11 in the women’s 200 backstroke, adding to the U.S. gold-medal haul.

The Australians claimed two gold medals Thursday. Grant Hackett won the 1,500 freestyle in a championship-record time of 14:41.65, and the women’s 4×100 medley relay also won gold.

It was Hackett’s fourth gold, along with the 800 freestyle and the 4×100 and 4×200 freestyle relays.

“I felt so smooth and comfortable,” Hackett said. “It’s a good feeling. … It normally hurts at 800 meters, but tonight it didn’t start hurting until 1,100 and I was surprised with the time.”